The origin of symmetrical flowers on both sides is one of the three key innovative events in the radiation differentiation process of angiosperms. Bilateral symmetric flowers often have horizontal orientation and asymmetric nectar conduction patterns, while radiative symmetric flowers typically have opposite sets of characteristics. Numerous studies in angiosperms, especially in core eudicots, have shown that the CYCLOIDEA (CYC) class genes of the TCP transcription factor family extensively determine the morphogenesis of bilaterally symmetric flowers. However, it is currently unclear how the floral orientation and nectar guidance modes were established.